A Saudi-led coalition said Sunday it launched a strike apparently against southern separatists in Yemen after they seized the presidential palace in the second city Aden.

The seizure, decried by the Riyadh-backed Yemeni government as a UAE-supported coup, reflects deep divisions between secessionists and loyalist forces, both of whom have fought Shiite Huthi rebels.

"The coalition targeted an area that poses a direct threat to one of the important sites of the legitimate government," a coalition statement said, calling on the separatist Southern Transitional Council to withdraw from positions seized in Aden or face further attacks.

It did not specify the target but residents in Aden told AFP it was an air strike against separatist camps in the city.

Riyadh-based Yemeni President Abedrabbo Mansour Hadi is backed by the coalition -- led by Saudi Arabia and its ally the United Arab Emirates -- that is battling the Iran-aligned Huthis.

But another force in the anti-Huthi coalition -- the UAE-trained Security Belt Force -- has since Wednesday been battling loyalists in Aden, the temporary base of Hadi's government.

The Security Belt Force is dominated by fighters who back the Southern Transitional Council (STC), which seeks to restore south Yemen as an independent state as it was from 1967-1990.

A Security Belt official told AFP Saturday the force had seized the presidential palace -- largely symbolic, due to Hadi's absence -- without a fight.

"Two hundred soldiers from the Presidential Guard were given safe passage out of the palace," the official said.

A witness confirmed the complex had been handed over.

The coalition called for an "immediate ceasefire", a spokesman was quoted as saying by the official Saudi Press Agency, and demanded an "urgent meeting" between the warring parties.

Saudi Deputy Defence Minister Khalid bin Salman also called for an "immediate cessation of hostilities and withdrawal from all forcefully occupied locations in Aden".

Both the Yemeni government and separatists said early Sunday they backed Riyadh's call for dialogue and a suspension of fighting.

- Government decries 'coup' -

Yemen's government earlier blamed the STC and the UAE for staging a "coup" against it.

The foreign ministry demanded "the UAE halt its material support and withdraw its military support, immediately and fully, from the groups that have rebelled against the state".

The STC's spokesman said the council was working to restore the water network, damaged in the fighting.

The International Crisis Group think tank warned that the latest clashes "threaten to tip southern Yemen into a civil war within a civil war."

"Such a conflict would deepen what is already the world's worst humanitarian crisis and make a national political settlement harder to achieve," it said.

Ties between the Security Belt and Hadi loyalists have been strained for years, and this week was not the first time they have engaged in armed clashes.

They fought a three-day battle in January 2018 that killed 38 people and wounded 222 others after the government prevented a rally by separatists.

The Security Belt has accused Hadi's backers of allowing Islamists into their ranks and of supporting the Muslim Brotherhood.

The Saudi-led coalition intervened in Yemen in 2015 to back the government against the Huthis, who are supported by Riyadh's regional rival Iran.

The Huthis had overrun large parts of northern and western Yemen, including the capital Sanaa, which they still control.

UAE Foreign Minister Abdullah bin Zayed Al-Nahyan said the UAE was "exerting all efforts to calm and de-escalate the situation", the official Emirati news agency WAM reported.

He added that the two camps should focus their efforts on fighting the Huthis instead of each other.

- 'Attacks on civilians' -

The latest violence flared on Wednesday during the funeral of a senior Security Belt commander killed earlier this month in a drone and missile attack on a training camp west of Aden.

The commander was among 36 people killed -- many of them newly graduated cadets -- in the aerial attack, claimed by the Huthis.

The UN human rights office later accused the Security Belt force of "reportedly carrying out and enabling retaliatory attacks against civilians" from northern Yemen.

The death toll is unclear, with initial reports saying 18 people, including civilians, had been killed on Thursday and Friday.

Doctors Without Borders (MSF) tweeted that it had treated more than 100 people in one of its hospitals in Aden.

Since 2015, fighting between the Huthis and Yemeni loyalists backed by the Saudi-led coalition has killed tens of thousands of people, mostly civilians, aid agencies say.

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